This is where biology meets demography: how populations grow, what keeps them in check, how species fit into communities, how those communities change over time, and how careful management can hold a habitat at the stage where biodiversity is highest.
Ecosystems are shaped by abiotic factors, biotic factors, and the niche each species occupies.
An ecosystem comprises a community of organisms together with the abiotic features of the environment with which they interact. The distribution of any species within an ecosystem is set where its tolerance to the abiotic factors and its competitive position against the biotic factors both allow it to persist.
Temperature sets enzyme rates and the metabolic ceiling. Light limits photosynthesis and primary production. pH affects enzyme activity and the availability of soil ions. Water availability and humidity govern cellular function and, in plants, transpiration.
Predation, competition, disease, mutualism, and parasitism. The living constraints on where a species can survive — interactions with other organisms that limit population size and distribution.
A niche is the full set of biotic and abiotic interactions a species engages in. A habitat is the place it lives. The two are not interchangeable.
Write
nichefor the ecological role andhabitatfor the place. AQA rejectshabitatas a substitute forniche.Write
communityfor the assemblage of species.Populationandecosystemare rejected as substitutes when the question asks for community.
Population size settles at the carrying capacity through intraspecific and interspecific competition.
Populations do not grow without limit. As numbers rise, resources per individual fall, intraspecific competition intensifies, and birth and death rates converge until the population stabilises near the carrying capacity — the environmental ceiling, not a y-axis label.
Write
carrying capacity. AQA rejectsmaximum capacity. The credited term names the environment as the limiting factor; the rejected substitute names only a number.
Intraspecific and interspecific competition compared.
| Competition type | Who competes | For what | Outcome |
|---|---|---|---|
| Intraspecific | Same species | Food, water, shelter, mates, light | Density-dependent regulation back toward carrying capacity |
| Interspecific | Different species | Shared resources where niches overlap | Less competitive species reduced or excluded |
Name the competition type explicitly:
intraspecificorinterspecific. The naming is itself a discrete mark in the mark scheme.AQA rejects
direct competition,allopatric competition, andspatial competitionas competition-type labels.
- Identify the competition type as interspecific — different species, shared resource.
- State that the more competitive species reduces the availability of the resource to the other.
- Trace the chain to the downstream molecular consequence: less light → less photosynthesis; less nitrate → less of a named nitrogen molecule (amino acid, protein, DNA, RNA, ATP); less phosphate → less of a named phosphorus molecule (DNA, RNA, ATP, phospholipid, RuBP, GP, TP, NADP).
- State the consequence at the population level: less growth, slower reproduction, smaller population.
End the mechanism chain at photosynthesis or at a named molecule. Stopping at
less lightorless nitratewithout naming what gets made less of forfeits the final mark — the molecular endpoint is what AQA credits.
The red and grey squirrel case is the standard worked example. Grey squirrels are larger and more generalist, extract more energy from acorns than red squirrels, and outcompete reds for food and nest sites where the two species share habitat. Red populations are reduced or eliminated in the overlap zones.
Predator and prey populations oscillate out of phase.
A predator captures and consumes a prey species; the two population sizes are coupled by a feedback loop with a lag, because a change in food availability takes time to translate into a change in predator birth rate.
The signature is the lag: on a graph, the predator peak follows the prey peak. The cycle is not the only mechanism regulating either population — disease, weather, and competition also act. In an answer, describe the feedback in directional terms (rises, falls, follows after a lag), not as a perpetual steady oscillation.
Sampling methods differ between sessile and mobile organisms.
Direct counting of every individual is rarely feasible. The technique is chosen by whether the organism moves. Sessile or slow-moving organisms — including all plants — are sampled by quadrat. Mobile animals are sampled by mark-release-recapture.
Plants are sampled by quadrat, not by mark-release-recapture. Plants cannot disperse after being marked, so marked individuals do not mix with the population and the Lincoln index assumption fails.
Quadrat and mark-release-recapture compared.
| Method | Applies to | Placement | Measure |
|---|---|---|---|
| Quadrat | Plants, sessile invertebrates | Random coordinates from a random number generator | Mean percentage cover or frequency, scaled up by area |
| Mark-release-recapture | Mobile animals | Second sample taken wherever marked individuals dispersed | Lincoln index from the two counts and the recaptured number |
Write
random number generatororrandom coordinates from a random number tablefor quadrat placement. AQA rejectsthrowing quadratsandrandomlywithout a named method.Write the calculation step in full: mean per quadrat multiplied by the number of quadrats in the total area.
Scale upalone is rejected.A precaution is what the investigator does (non-toxic mark, mark invisible to predators). An assumption is a condition that must hold (closed population, equal probability of capture). They are graded separately.
N = (first catch × second catch) ÷ number of marked recaptures, where N is the estimated population size. The formula assumes the proportion of marks in the second sample equals the proportion of marks in the whole population.
- Capture a first sample, mark each individual in a way that does not affect survival or detectability, and release them into the same area.
- Wait long enough for the marked individuals to disperse and mix randomly with the unmarked population.
- Capture a second sample, count how many carry the mark, and apply the Lincoln index.
Closed population (no significant immigration, emigration, births, or deaths during the study). Random mixing of marked and unmarked individuals before the second sample. Mark does not affect survival or detectability. Mark persists until the second sampling.
Write
at least 10 quadratsorlarge sample. AQA rejectsseveralas a sample-size description.
Pitfall — The three-mark sampling checklist
Every quadrat or MRR question allocates three independent marks. Drop any one and the answer caps below maximum.
- Named random placement method — a random number generator or random table, not "randomly" without a named method.
- Large sample size — at least 10 quadrats, or a stated number, not "several".
- The calculation step in full — mean per quadrat × number of quadrats in the total area, or the Lincoln index written out. "Scale up" alone forfeits this mark.
The calculation step is the most consistently dropped. The answer is not complete until the multiplication or division is written.
Succession is a sequence of communities that change their own environment.
Succession is the directional replacement of one community by another over time. The mechanism is species-driven environmental change — each community modifies the abiotic conditions in ways that allow the next community to establish and that make conditions less suitable for itself.
- Name a pioneer species from the figure (lichens or mosses on bare rock, for primary succession).
- State an abiotic change with a named mechanism: forms soil; adds humus; stabilises the substrate; increases water-holding capacity.
- State the consequence using the credited phrase: conditions are less hostile or more suitable for the next species.
- Name the climax community from the figure — the stable end-point under the prevailing climate (woodland in the temperate UK).
Write
pioneer species. AQA rejectsprimary speciesandcoloniser.Write
climax communityin full.Climaxalone is not credited.Write
less hostileormore suitable. AQA rejectsmore habitableandmore hospitableas substitutes.
Primary succession begins on bare substrate with no pre-existing soil — newly exposed rock or coastal sand. Secondary succession begins on previously inhabited ground where soil is already present, so it bypasses the early seral stages and proceeds more rapidly.
On a succession graph, time runs left to right: pioneer at the left, climax at the right. Reading the axis backwards — calling the trees pioneers and the grasses the climax — is the single most common error on this question type.
A seral stage is one community in the sequence. The whole sequence is a sere. The climax community is the stable end-point whose composition does not change further under the prevailing climate.
Conservation often holds a habitat at an earlier seral stage.
Many habitats of high conservation value — heathland, moorland, chalk grassland, reed beds — are not climax communities. Left unmanaged, each would progress through succession to its local climax, and the species characteristic of the open earlier-stage habitat would be lost. Conservation management arrests or resets the sequence.
Burning removes the above-ground biomass of shrubs and small trees, eliminating the species characteristic of later seral stages. Root systems, seed banks, and the soil layer are preserved, so the habitat re-establishes from an early seral stage rather than from bare rock. Periodic burning prevents climax and holds biodiversity.
- Sterile males of the pest species are mass-released into the wild population and compete with fertile males for mates. The named mechanism is intraspecific competition.
- Matings with sterile males produce no viable offspring.
- Birth rate falls and the population declines over successive generations.
The sterile insect mechanism is intraspecific competition followed by failed reproduction. AQA rejects answers that explain the technique via the sterile males' inability to transmit disease — female mosquitoes, not males, bite, and the technique is about reproduction, not pathogenicity.
When sterile insect release fails or underperforms, the credited explanation is a fitness effect of the sterilising radiation — reduced lifespan, reduced courtship success, reduced attractiveness to females. Disease-transmission answers are rejected here too.
Key terms
- carrying capacity
- intraspecific competition
- interspecific competition
- niche
- quadrat
- mark-release-recapture
- Lincoln index
- succession
- pioneer species
- climax community